Abundance-based Coverage Estimator (ACE) of total species richness (Chao & Lee 1992). Separates species into rare and abundant groups based on an abundance threshold.
Details
ACE partitions species into rare (abundance \(\le\) threshold) and abundant (abundance > threshold) groups. The estimate is: $$S_{ACE} = S_{abund} + \frac{S_{rare}}{C_{ACE}} + \frac{f_1}{C_{ACE}} \gamma^2$$ where \(C_{ACE}\) is the sample coverage of rare species and \(\gamma^2\) is the estimated coefficient of variation.